Four plant species have been described from the Bodie Hills. Some kind of synchronicity must have been in effect, because all four were collected in the summer of 1945. Three of them are mustards (Brassicaceae), and all of those were described by Reed C. Rollins (a professor at Harvard University, and a renowned expert in the taxonomy of the mustard family). Two of the type specimens were collected by Annie Alexander and Louise Kellogg (ambitious collectors of the California flora from 1939 to 1949); the other two were collected — on the same day in the same location, perhaps only minutes apart — by Ira Wiggins (a professor at Stanford and author of several floras) together with Reed Rollins. All four of these plants have a similar, very limited distribution in northern Mono, southern Lyon, and western Mineral counties, with the majority of known occurrences being in the Bodie Hills.
Published descriptions of these and all other plant species reference a type specimen and a type locality. A type specimen is an individual specimen (or a group of specimens) to which a scientific name is formally attached. For vascular plants, this is usually an 11.5 x 16.5-inch sheet of herbarium paper with a pressed specimen and one or more printed labels glued onto it. The collector’s collection number and the herbarium’s accession number are on the sheet so the exact same specimen can be found and examined again later. A type locality is the geographical location where the type specimen was originally found.
Here are the four plants with type localities in the Bodie Hills:
Boechera bodiensis (Bodie Hills rock-cress) was described by Reed Rollins (in Contributions of the Gray Herbarium 212:113, 1982) from material originally identified (in 1945) as a hybrid of Arabis sparsiflora and A. fernaldiana. Additional specimens collected over the next 4 decades provided the basis for its recognition as a new species in 1982. In the early 2000s, molecular studies showed that Arabis actually consisted of two distantly related clades, with morphological similarities attributed to evolutionary convergence. All the species in the Bodie Hills previously treated as Arabis now belong in Boechera. Boechera species are notoriously difficult to define, key, and identify. The Flora of North America notes that “a rare confluence of hybridization, apomixis, and polyploidy makes this one of the most difficult genera in the North American flora.” Perhaps it’s still an actively evolving group. Boechera bodiensis is still regarded as being of hybrid origin, but with B. falcifructa as one of the parents.
The type specimen is Ira L. Wiggins and Reed C. Rollins #536, collected on August 3, 1945 (UC727326) northwest of Masonic Peak, perhaps between Chemung Mine and Lakeview Spring. Habitats of Bodie Hills rock-cress include dry, open, rocky, high or north-facing slopes, exposed rocky ridges and summits, moisture-accumulating microsites in sagebrush, under shrubs, and disturbed soils of prospector’s diggings.
Boechera bodiensis has been found mostly on and around Masonic Mountain in the Bodie Hills and Glass Mountain, southeast of Mono Lake. A few additional collections are from the Wassuk Range and the southern White Mountains.
Streptanthus oliganthus (Masonic Mountain jewelflower) was described by Reed Rollins (in Contributions of the Dudley Herbarium 3(11):372-373, 1946) from another collection by Wiggins and Rollins on the same day (maybe even the same time and location) as the type specimen for Boechera bodiensis. The type specimen is Ira L. Wiggins and Reed C. Rollins #535, collected on August 3, 1945 (UC727392) (see the specimen here).
Streptanthus oliganthus grows in dry, open pinyon pine woodland and sagebrush scrub habitat. Most collections have been around Masonic Mountain and the east side of the Sweetwater Mountains. Collections near Sonora Pass, in the White Mountains near Westgard Pass, and in the Copper Mountain area southwest of Conway Summit have been attributed to S. oliganthus, but may be S. cordatus. Streptanthus cordatus is similar in size and habitat, but is is more prevalent in the eastern Sierra Nevada and Great Basin ranges. The two are distinguished as follows:
Cusickiella quadricostata (Bodie Hills Cusickiella) was described by Reed Rollins (in Contributions of the Dudley Herbarium 3:366, 1946) as Draba quadricostata. Cusickiella (named for W.C. Cusick, an Oregon plant collector, 1842–1922) is a small group of only two species now segregated from Draba. Cusickiella differs from Draba most noticablely in the shape of the fruits. The fruits of Cusickiella have rounded or keeled valves, whereas in Draba the valves are typically cylindric or flat. Also, the fruits of Cusickiella contain only 1–4 ovules or seeds, whereas Draba has 10 or more.
The type specimen of Cusickiella quadricostata is Annie Alexander & Louise Kellogg #4543 (UC694166), collected on July 28, 1945. The type locality is “on the road to Bodie, 2 miles southwest of Masonic Spring, southeast flank of Masonic Mountain, altitude 8600 feet.” They encountered this plant earlier the same day at New York Hill (#4540), and might have found it around the north side of Masonic Mountain too, had they stopped in the right place. This specimen (#4543) appears to be the very last collection the team of Alexander and Kellogg ever made in the Bodie Hills. They collected elsewhere in Mono County in 1946, including in Bridgeport Meadows, but they did not return to the Bodie Hills.
Cusickiella quadricostata is known from quite a few locations in the Bodie Hills, with additional locations in the Sweetwater Mountains, Pine Grove Hills, and southern Wassuk Range of Mono, Mineral, and Lyon counties. Its habitat is usually gravelly slopes, ridges, and flats, associated with scattered low sagebrush or cusion plants. The other species of Cusickiella, C. douglasii, is also found in the Bodie Hills, but it has a much wider range, extending to Nevada, Washington, Idaho, and Utah. They differ as follows:
Phacelia monoensis (Mono County phacelia) was described by Richard Halse, of Oregon State University in Corvallis (Madroño 28:124, 1981) from material previously identified as Miltitsia lutea. Miltitzia is a group of yellow-flowered annuals now treated as a section of Phacelia, in the part of Boraginaceae previously treated as Hydrophyllaceae. The type specimen is Annie Alexander & Louise Kellogg #4346 (UC736041), collected on June 30 1945.
The type locality, quoted from the specimen label, is “Altitude 7375 feet; in scraped ground of red, caked adobe above road and meadow, Mormon Ranch, 8.5 miles south-west of Bodie.” The label also notes that the plants were “associated with Nemacladus rigidus.” Topographic maps of this area from 1911 to 1958 place “Mormon Ranch” near the east end of Mormon Meadow, about where Clearwater Creek crosses today’s State Route 270. The nearest and most extensive area of clayey red soil is on the low hills on the south side of Mormon Meadow, just east of today’s Coyote Springs Road. Unfortunately, much of this area has been heavily trampled for several decades by sheep concentrated around a sheep herder’s camp.
Phacelia monoensis is known from several other locations in the Bodie Hills, Sweetwater Mountains, Pine Grove Hills, and fer-northern White Mountains in Mono, Mineral, Lyon, and perhaps Esmeralda counties. The “monoensis” epithet is apt, because a majority of the known populations are still to be found in Mono County. It favors dark red or red-brown clayey soils that are loosened by natural shrink-swell processes or by the occasional passing of vehicles along unpaved roads.
Bonus: a Mineral, Bodieite
Minerals have type specimens and type localities too. The recently-described mineral Bodieite has one of its two co-type localities in the Bodie Hills near Masonic Mountain.
Bodieite (photos here) is a soft, colorless to yellow or green, crystalline mineral. It is unique in being both a tellurate and a sulfate of bismuth [Bi2(TeO3)2(SO4)]. Bodieite was “named for the Bodie Hills volcanic field, in which the Pittsburg-Liberty mine is located, and for the town of Bodie, California, which is about 19 km SSE of the Pittsburg-Liberty mine.”
Bodieite has two rather widely separated co-type localities: (1) the Pittsburg-Liberty Mine, at New York Hill in the Masonic District of the northern Bodie Hills, and (2) tailings of the North Star Mine (Star Consolidated Mine), on the south side of Mammoth Peak, near Mammoth (but not the Mammoth in Mono County), in the East Tintic Mountains of Juab County, Utah (southwest of Provo and Utah Lake).
Thanks to Jim Morefield and Janel Johnson for their photos of the Boechera and the Streptanthus!
Copyright © Tim Messick 2019. All rights reserved.
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