Category Archives: Plant Identification

Plants Described from the Bodie Hills

Masonic Mountain from the north

Masonic Mountain from the north. Three plant species were described from here.

Four plant species have been described from the Bodie Hills. Some kind of synchronicity must have been in effect, because all four were collected in the summer of 1945. Three of them are mustards (Brassicaceae), and all of those were described by Reed C. Rollins (a professor at Harvard University, and a renowned expert in the taxonomy of the mustard family). Two of the type specimens were collected by Annie Alexander  and Louise Kellogg (ambitious collectors of the California flora from 1939 to 1949); the other two were collected — on the same day in the same location, perhaps only minutes apart — by Ira Wiggins (a professor at Stanford and author of several floras) together with Reed Rollins. All four of these plants have a similar, very limited distribution in northern Mono, southern Lyon, and western Mineral counties, with the majority of known occurrences being in the Bodie Hills.

Published descriptions of these and all other plant species reference a type specimen and a type locality. A type specimen is an individual specimen (or a group of specimens) to which a scientific name is formally attached. For vascular plants, this is usually an 11.5 x 16.5-inch sheet of herbarium paper with a pressed specimen and one or more printed labels glued onto it. The collector’s collection number and the herbarium’s accession number are on the sheet so the exact same specimen can be found and examined again later. A type locality is the geographical location where the type specimen was originally found.

Here are the four plants with type localities in the Bodie Hills:


Boechera bodiensis (Bodie Hills rock-cress) was described by Reed Rollins (in Contributions of the Gray Herbarium 212:113, 1982) from material originally identified (in 1945) as a hybrid of Arabis sparsiflora and A. fernaldiana. Additional specimens collected over the next 4 decades provided the basis for its recognition as a new species in 1982. In the early 2000s, molecular studies showed that Arabis actually consisted of two distantly related clades, with morphological similarities attributed to evolutionary convergence. All the species in the Bodie Hills previously treated as Arabis now belong in Boechera. Boechera species are notoriously difficult to define, key, and identify. The Flora of North America notes that “a rare confluence of hybridization, apomixis, and polyploidy makes this one of the most difficult genera in the North American flora.” Perhaps it’s still an actively evolving group. Boechera bodiensis is still regarded as being of hybrid origin, but with B. falcifructa as one of the parents.

Boechera bodiensis

Boechera bodiensis with sagebrush (Photo © James D. Morefield via Natureserve)

The type specimen is Ira L. Wiggins and Reed C. Rollins #536, collected on August 3, 1945 (UC727326) northwest of Masonic Peak, perhaps between Chemung Mine and Lakeview Spring. Habitats of Bodie Hills rock-cress include dry, open, rocky, high or north-facing slopes, exposed rocky ridges and summits, moisture-accumulating microsites in sagebrush, under shrubs, and disturbed soils of prospector’s diggings.

Boechera bodiensis has been found mostly on and around Masonic Mountain in the Bodie Hills and Glass Mountain, southeast of Mono Lake. A few additional collections are from the Wassuk Range and the southern White Mountains.

Boechera bodiensis

Boechera bodiensis in flower (Photo © James D. Morefield)


Streptanthus oliganthus (Masonic Mountain jewelflower) was described by Reed Rollins (in Contributions of the Dudley Herbarium 3(11):372-373, 1946) from another collection by Wiggins and Rollins on the same day (maybe even the same time and location) as the type specimen for Boechera bodiensis. The type specimen is Ira L. Wiggins and Reed C. Rollins #535, collected on August 3, 1945 (UC727392) (see the specimen here).

Streptanthus oliganthus

Streptanthus oliganthus (Photo © Janel Johnson via iNaturalist, CC BY-NC 4.0)

Streptanthus oliganthus grows in dry, open pinyon pine woodland and sagebrush scrub habitat. Most collections have been around Masonic Mountain and the east side of the Sweetwater Mountains. Collections near Sonora Pass, in the White Mountains near Westgard Pass, and in the Copper Mountain area southwest of Conway Summit have been attributed to S. oliganthus, but may be S. cordatus. Streptanthus cordatus is similar in size and habitat, but is is more prevalent in the eastern Sierra Nevada and Great Basin ranges. The two are  distinguished as follows:

Streptanthus key


Cusickiella quadricostata (Bodie Hills Cusickiella) was described by Reed Rollins (in Contributions of the Dudley Herbarium 3:366, 1946) as Draba quadricostata. Cusickiella (named for W.C. Cusick, an Oregon plant collector, 1842–1922) is a small group of only two species now segregated from Draba. Cusickiella differs from Draba most noticablely in the shape of the fruits. The fruits of Cusickiella have rounded or keeled valves, whereas in Draba the valves are typically cylindric or flat. Also, the fruits of Cusickiella contain only 1–4 ovules or seeds, whereas Draba has 10 or more.

Cusickiella quadricostata

Cusickiella quadricostata (Photo © Tim Messick)

The type specimen of Cusickiella quadricostata is Annie Alexander & Louise Kellogg #4543 (UC694166), collected on July 28, 1945. The type locality is “on the road to Bodie, 2 miles southwest of Masonic Spring, southeast flank of Masonic Mountain, altitude 8600 feet.” They encountered this plant earlier the same day at New York Hill (#4540), and might have found it around the north side of Masonic Mountain too, had they stopped in the right place. This specimen (#4543) appears to be the very last collection the team of Alexander and Kellogg ever made in the Bodie Hills. They collected elsewhere in Mono County in 1946, including in Bridgeport Meadows, but they did not return to the Bodie Hills.

Cusickiella quadricostata is known from quite a few locations in the Bodie Hills, with additional locations in the Sweetwater Mountains, Pine Grove Hills, and southern Wassuk Range of Mono, Mineral, and Lyon counties. Its habitat is usually gravelly slopes, ridges, and flats, associated with scattered low sagebrush or cusion plants. The other species of Cusickiella, C. douglasii, is also found in the Bodie Hills, but it has a much wider range, extending to Nevada, Washington, Idaho, and Utah. They differ as follows:

Cusickiella key


Phacelia monoensis (Mono County phacelia) was described by Richard Halse, of Oregon State University in Corvallis (Madroño 28:124, 1981) from material previously identified as Miltitsia lutea. Miltitzia is a group of yellow-flowered annuals now treated as a section of Phacelia, in the part of Boraginaceae previously treated as Hydrophyllaceae. The type specimen is Annie Alexander & Louise Kellogg #4346 (UC736041), collected on June 30 1945.

Phacelia monoensis

Phacelia monoensis (Photo © Tim Messick)

The type locality, quoted from the specimen label, is “Altitude 7375 feet; in scraped ground of red, caked adobe above road and meadow, Mormon Ranch, 8.5 miles south-west of Bodie.” The label also notes that the plants were “associated with Nemacladus rigidus.” Topographic maps of this area from 1911 to 1958 place “Mormon Ranch” near the east end of Mormon Meadow, about where Clearwater Creek crosses today’s State Route 270. The nearest and most extensive area of clayey red soil is on the low hills on the south side of Mormon Meadow, just east of today’s Coyote Springs Road. Unfortunately, much of this area has been heavily trampled for several decades by sheep concentrated around a sheep herder’s camp.

Phacelia monoensis is known from several other locations in the Bodie Hills, Sweetwater Mountains, Pine Grove Hills, and fer-northern White Mountains in Mono, Mineral, Lyon, and perhaps Esmeralda counties. The “monoensis” epithet is apt, because a majority of the known populations are still to be found in Mono County. It favors dark red or red-brown clayey soils that are loosened by natural shrink-swell processes or by the occasional passing of vehicles along unpaved roads.

Phacelia monoensis

Phacelia monoensis (Photo © Tim Messick)


Bonus: a Mineral, Bodieite
Minerals have type specimens and type localities too. The recently-described mineral Bodieite has one of its two co-type localities in the Bodie Hills near Masonic Mountain.

Bodieite (photos here) is a soft, colorless to yellow or green, crystalline mineral. It is unique in being both a tellurate and a sulfate of bismuth [Bi2(TeO3)2(SO4)]. Bodieite was “named for the Bodie Hills volcanic field, in which the Pittsburg-Liberty mine is located, and for the town of Bodie, California, which is about 19 km SSE of the Pittsburg-Liberty mine.”

Bodieite has two rather widely separated co-type localities: (1) the Pittsburg-Liberty Mine, at New York Hill in the Masonic District of the northern Bodie Hills, and (2) tailings of the North Star Mine (Star Consolidated Mine), on the south side of Mammoth Peak, near Mammoth (but not the Mammoth in Mono County), in the East Tintic Mountains of Juab County, Utah (southwest of Provo and Utah Lake).


Thanks to Jim Morefield and Janel Johnson for their photos of the Boechera and the Streptanthus!


Copyright © Tim Messick 2019. All rights reserved.
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Bailey’s Greasewood in the Bodie Hills

Sarcobatus baileyi

Sarcobatus baileyi (note the short, pubescent, clustered leaves)

Floristic studies of large areas are always works in progress. New things regularly turn up in previously unexplored habitats or in more favorable water years. Even reviewing photographs taken several years ago can reveal “new finds” among plants previously labeled with a big question mark.

Just recently, looking through my Lightroom catalog at photos taken 3 and 4 years ago, I realized I had photographed Bailey’s greasewood (Sarcobatus baileyi), a plant I was unaware of at the time, but have since come to know a bit through observations by others on iNaturalist. I had noticed, photographed, and scratched up my hands on it twice: in May 2015, on a ridge just south of the Sweetwater-Aurora road (NF-028) and west of Red Wash Creek (in Lyon County, Nevada); then in September 2016 several miles farther southeast, where NF-028 crosses a low ridge to enter the west end of Fletcher Valley (in Mineral County). Only in 2019 have I recognized what it was, but I think it may be locally somewhat common on gravelly ridges in the low northern and eastern foothills on the Nevada side of the Bodie Hills.

Sarcobatus baileyi

Sarcobatus baileyi

The growth form, foliage, and habitat of Sarcobatus baileyi tend to be so different from the much more common and familiar S. vermiculatus, that it scarcely registers as a greasewood if you aren’t aware it exists. The flowering parts (male and female in separate clusters, and scarcely resembling “flowers” at first glance) are similar in both species, although they are nearly sessile in S. baileyi and are on longer lateral branches in S. vermiculatus. Here’s a key to Sarcobatous species, adapted from a few different sources:

1. Plants generally erect (infrequently ± spheric), often 1–2 m tall, loosely and irregularly branched, the lowest branches rarely in contact with the ground; leaves 1–4 cm, solitary on elongate shoots of current season, usually glabrous (may be slightly pubescent); pistillate flowers and staminate spikes on long lateral branches with 3–9 obvious internodes; mature staminate spikes generally 10–40 mm; fruit 2.5–5.2 mm long, the wing 4–10 mm wide; plants common, usually associated with seasonally moist, saline or alkaline soils, often phreatophytic (rooting deeply to reach the water table)
. . . . . . . . . . . . . . . . . . . . .   S. vermiculatus

1′ Plants generally ± spheric, low, < 1 m tall, densely and intricately branched, the lowest branches in contact with the ground; leaves shorter, 0.5–1.5 cm, clustered in short shoots on cushion-like pads of longer or older twigs, usually pubescent; pistillate flowers and staminate spikes appearing ± sessile, but on short branches with 1–3 minute internodes; mature staminate spikes generally < 10 mm; fruit 5–11.5 mm long, the wing 7–16 mm wide; plants uncommon, frequently on soils that are not obviously saline or alkaline, commonly non-phreatophytic (i.e., shallower-rooted)
. . . . . . . . . . . . . . . . . . . . .   S. baileyi

Sarcobatus baileyi habitat

Sarcobatus baileyi habitat (they are the greenest low shrubs here)

Bailey’s greasewood is distributed across the central and western Great Basin, nearly all in Nevada. Bailey’s greasewood has been collected in California east of the White Mountains in Fish Lake Valley, along Hwy 266, southeast of Dyer, Nevada, just barely inside Mono County. Calflora indicates it has been observed near Honey Lake in Lassen County and a paper on Sacrobatus ecology (Drenovsky et al. 2011) documents its presence south of Owens Lake in Inyo County.

Common greasewood (S. vermiculatus) has a much wider distribution across all of the Great basin and beyond, to Canada, the Great Plains, and Mexico. In and near the Bodie Hills, it’s common at Travertine Hot Springs, around Mono Lake, other moist alkaline meadow margins, and rarely in uplands or seasonal washes.

Sarcobatus vermiculatus

Sarcobatus vermiculatus (with longer leaves, not clustered)

Sarcobatus vermiculatus

Sarcobatus vermiculatus (with a Variegated meadowhawk, Sympetrum corruptum)

Sarcobatus baileyi and S. vermiculatus sometimes grow together and they sometimes intergrade; because of this, some individuals may not be clearly assignable to one species or the other. Intergradation may result from hybridization, soil or moisture stress, or in some cases pathogens or insect activity. Such intergradation has not been observed in the Bodie Hills, however. With somewhat limited habitat for both species, they probably remain fairly distinct here.

Vernon O Bailey

Vernon O. Bailey on horseback in Nevada, 1898. (Source: Smithsonian Institution Archives, public domain)

Who, then, was Bailey?
Vernon O. Bailey (1864–1942) was an American naturalist, mammalogist with the Death Valley Expedition of 1890-1891, and a Chief Field Naturalist for the Bureau of Biological Survey (part of what became the U.S. Fish and Wildlife Service). His name is commemorated in Canis lupus baileyi, the Mexican wolf, and several other species of plants (including species of Chrysothamnus, Echinocereus, and Yucca). (Other western plants, including an Eriogonum, have been named for at least one other Bailey—William Whitman Bailey.) The Sarcobatus was described by Frederick Vernon Coville, an American botanist (1867–1937) who was also in the Death Valley Expedition and wrote the report on its botanical findings. Later, Coville was chief botanist at the U.S. Department of Agriculture and first director of the U.S. National Arboretum.

In Coville’s 1892 description of Sarcobatus baileyi, he says, “The plant was first seen by Mr. [Vernon O.] Bailey at Cloverdale, Esmeralda County, Nevada, in 1890, and recognized by him as different from S. vermiculatus. In company with Dr. [Clinton Hart] Merriam he afterwards found it in a valley in Nye County, Nevada, southeast by east from Gold Mountain, near Thorpe’s quartz mill, and later in Fish Lake Valley … on the California State line. … I take pleasure in associating Mr. Bailey’s name with this shrub, both as a mark of his earnest and invaluable labors in the field of natural history and as a reminder of a warm friendship established among the vicissitudes of a desert exploration.” Vernon Bailey and C. Hart Merriam were not only close professional colleagues; they also became in-laws: in 1899 Bailey married Merriam’s sister—and prominent ornithologist—Florence Merriam.


Copyright © Tim Messick 2019. All rights reserved.
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Plants of the Bodie Hills Checklist: January 2019 Edition

Plants of the Bodie Hills Checklist Cover 2019

Another year has passed and a surprising number of taxa (species, varieties, and subspecies) have been added to Plants of the Bodie Hills: an Annotated Checklist — about 50! Some of these are new finds in the field; some are new records in the on-line herbarium databases. Some of these are definite new additions; some are in the “maybe” category. The total count is now about 750 taxa.

Other changes in this year’s edition include the addition of keys to selected genera, and additions to the map on the last page (or back cover, if it’s printed 2-sided).

CLICK HERE to visit the Downloads page. The January 2019 edition of the checklist is a 64-page, 6.1 mb PDF file.

Here’s a selection of favorite plants seen during visits to the Bodie Hills in 2018:

Eriogonum caespitosum

Matted wild buckwheat: Eriogonum caespitosum

Eriogonum maculatum

Spotted wild buckwheat: Eriogonum maculatum

Salvia dorrii

Dorr’s sage: Salvia dorrii

Thelypodium laciniatum

Feathery thelypodium: Thelypodium laciniatum

Allium anceps

Twin leaved onion: Allium anceps

Angelica capitellata

Ranger’s buttons or Swamp white heads: Angelica capitellata
with lots of visiting insects.

Astragalus curvicarpus

Curvepod milkvetch: Astragalus curvicarpus

Rosa woodsii

Woods rose: Rosa woodsii subsp. ultramontana


Copyright © Tim Messick 2019. All rights reserved.
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