Tag Archives: Mono County

Plants of the Bodie Hills, 2025 Edition

Plants of the Bodie Hills, 2025 Edition, is now available on the Downloads page (a free PDF). As in previous years, the new edition contains additions, corrections, nomenclatural updates, and refinements to the keys.

Nearly 700 “minimum rank” taxa in 79 families are known to occur in the Bodie Hills. (Minimum rank taxa are the subspecies or varieties within a species, if the species has those, or just the species itself if it contains no lower taxa.) Up to about 100 more such taxa may be possible here—because they occur nearby and suitable habitats are present, or because they were collected very long ago but have not been seen in recent decades. These remain uncertain, not yet confirmed by recent collections or observations within the Bodie Hills.


Here are the notable changes included in this year’s edition:

Two vascular plants not previously known or expected to occur in the Bodie Hills were recently documented here:

Erigeron linearis (Desert yellow fleabane) was observed in 2023 in talus on a high ridge west of Masonic Mountain and in 2024 at two locations on rocky slopes in sagebrush scrub south of Hwy 270, east of Mormon Meadow (https://www.inaturalist.org/observations?captive=false&place_id=121648&subview=map&taxon_id=76926).

Erigeron linearis on Masonic Mountain (by Chloe and Trevor Van Loon, on iNaturalist)

A Parnassia (Grass-of-Parnassus), probably P. palustris, was discovered in late July 2024 at a hillside spring near the east end of Cow Camp Road. The observation photos were somewhat inconclusive as to whether this was P. palustris or P. parviflora, so follow-up site visits (in July–August) are needed to confirm which one is present.

Parnassia sp. (by Mike Heine, on iNaturalist)

Several plants that had been previously reported or expected to occur in the Bodie Hills were confirmed to be present:

Allium lemmonii (Lemmon’s onion) was previously observed in 2023 and confirmed in 2024 near Success Mine and at Chemung Lake. This is typically on seasonally saturated clayey soils. As with many wild onions, one has to closely examine the shape of the ovary crests (around the center of the flower) to identify the species. In Allium lemmonii, the crests are thick and rounded in cross-section; in other species that may appear similar (A. atrorubens and A. bisceptrum), the ovary crests are thin and triangular.

Allium lemmonii (note the thick ovary crests)

Tiquilia nuttallii (Annual tiquilia) is common in sandy soils throughout the Mono Basin, and as expected, has been found along the southwestern and northeastern edges of the Bodie Hills. The genus Tiquilia was formerly in the broadly-defined Boraginaceae (Borage or Forget-me-not family), but under the newer 11-family concept of order Boraginales, supported by much genetic work, Tiquilia is now in Ehretiaceae (Ehretia family).

Tiquilia nuttallii

Geranium californicum (California geranium) was collected at or near Bodie Ranch (5.3 road miles northeast of Bodie) by Annie Alexander and Louise Kellogg in 1945 and has not been collected or observed in the Bodie Hills since then. In May of 2024, a Geranium assumed to be G. californicum was discovered near this location—in leaf, but not yet in flower. Flower characters rather than leaf characters are necessary for distinguishing G. californicum from G. richardsonii, another common species of the Sierra Nevada. This new observation is most likely the same species observed by Alexander and Kellogg, but further observations of the flowers are needed to confirm which of these two potential species it is.

An often-misidentified (by me and others) lupine along the Bodie-Masonic road near Bodie Mountain was confirmed to be Lupinus angustiflorus (Narrowflower lupine), not L. andersonii (Anderson’s lupine). Vegetatively they are quite similar, and are distinguished only by subtle characters of the flower and inflorescence. In Lupinus angustiflorus the banner back is generally hairy (but the hairs may be very sparse), the banner spot is orange to yellow, and the inflorescence bracts are more or less persistent. In Lupinus andersonii the banner back is glabrous, the banner spot is white turning purple, and the inflorescence bracts are deciduous. These lupines are best keyed with fresh flowers. The hairiness of banner backs is often best seen on flowers near the top of the inflorescence that are still in bud.

Lupinus angustiflorus near Bodie Mountain

The small and easily overlooked Suaeda occidentalis (Western seablite), which I had collected near Travertine Hot Springs in 1981, was again found there in 2024. It occurs on moist or dried, alkaline soils and is uncommon in California, but more common throughout the Great Basin.

Suaeda occidentalis at Travertine Hot Springs

Three more species are likely additions, but need confirmation:

  • Sphaeralcea grossulariifolia (Gooseberry-leaved globemallow) was collected on the west side of the east Walker River along Highway 182—so, technically on the very edge of the Sweetwater Mountains—but if it’s there, then it’s also likely to occur somewhere in the northern Bodie Hills.
  • An immature (not yet flowering) specimen that appears to be Solidago multiradiata (Northern goldenrod) was found at Murphy Spring. Let’s hope this can be confirmed in the future with plants in flower.
  • An Agoseris (False dandelion) that might be A. retrorsa (Spearleaf Agoseris) was observed in 2023 in the southwestern Bodie Hills at Sage Flat.

Two “mystery plants” remain to be properly identified: an Aphyllon (Broomrape) along the road to Aurora from Fletcher, and a Pinus (likely P. flexilis, Limber pine) at the top of a steep slope near Millersville. The Aphyllon beside the road to Aurora may be the same as one observed in 2022 but posted just recently to iNaturalist from Fletcher Valley, near Nine-Mile Ranch.


Aside from these species additions and confirmations, the 2025 edition includes substantially revised or improved keys to species in several genera, notably Arnica (Arnicas), Chrysothamnus (Rabbitbrushes), Solidago (Goldenrods), and Salix (Willows).

Finally, a major new feature this year is an index to the scientific and common names of families and genera. In the PDF, the page numbers in the index are live links, so you can click (or tap) on those to immediately go to the indicated page. The many other web links throughout the document will open the indicated resource in your web browser.

As before, you have two options for how to use this document: 1) load the PDF onto a mobile device or 2) print the PDF yourself.

  • Using a mobile device: I’ve found the PDF to be quite readable on my iPhone (in the Books app), although it helps that I’m near-sighted. It’s even easier to read on a tablet or laptop.
  • Printing the PDF: You can print the PDF yourself or at a local print shop. I highly recommend printing the 136 pages 2-sided to conserve paper and to reduce bulk and weight in the field. A comb or spiral binding, binder clip, or other binding will hold it together.

Thanks to everyone who posts their observations from the Bodie Hills to iNaturalist. This citizen-science platform has provided much useful information for Plants of the Bodie Hills over the last several years. Thanks also to herbaria that post recent collections to online databases. And special thanks this year to Chauncey Parker for his painstaking proofreading of the manuscript (but any remaining errors are mine).

Your additions, corrections, comments, or questions are always welcome.

September snow on Potato Peak

Copyright © Tim Messick 2025. All rights reserved.

Plants of the Bodie Hills, 2024 Edition

Plants of the Bodie Hills, 2024 Edition, is now available on the Downloads page (a free PDF). As in previous years, the new edition contains additions, corrections, nomenclatural updates, and refinements to the keys. A key to families of Dicots (Magnoliopsida) in the area has at long last been added. Several place names, road names, and boundaries have been added to the map on the back cover (last page of the PDF).

The total number of “minimum rank” vascular plant taxa documented to occur in the Bodie Hills now stands at 727, in 80 families. Another 96 taxa, including 2 additional dicot families, are still considered to be potentially present, but not yet confirmed by collections or observations within the Bodie Hills.

Below Travertine Hot Springs

New additions to the 2024 edition (found during 2023) are:

  • Lupinus uncialis (Lilliput lupine) at the Bodie Hills Preserve and at Travertine Hot Springs,
  • Micromonolepis pusilla (Dwarf monolepis) at Mormon Meadow,
  • Phacelia curvipes (Washoe phacelia) in the southwestern Bodie Hills,
  • Platanthera tescamnis (Great Basin bog orchid) at a remote spring in the northeastern Bodie Hills in Mineral County, and
  • Saponaria officinalis (Common soapwort) at Conway Summit.

Plants seen in 2023 that had been expected in the Bodie Hills or not seen here for decades were:

  • Antirrhinum (Sairocarpus) kingii (King’s snapdragon) at Travertine Hot Springs,
  • Collomia grandiflora (Large-flowered collomia) near Mormon Meadow,
  • Eriogonum hookeri (Hooker’s buckwheat) at Travertine Hot Springs,
  • Erythranthe rubella (Little redstem monkeyflower) at “Sage Flat,” (near the Little Bodie Mine historical marker),
  • Malacothrix glabrata (Desert dandelion) in Red Creek Wash (from a 1983 collection),
  • Penstemon deustus (Hot rock penstemon) north of Bridgeport Reservoir Dam, and
  • Silene nuda (Sticky catchfly) along Cow Camp Road.

Nearly all of these were found, not by me, but by others posting their observations to iNaturalist (thanks to all who do this!). All observations within the Bodie Hills on iNaturalist can be seen HERE.

Jeffrey and Lodgepole pines in the western Bodie Hills

As before, you have two options for how to use this document: 1) load the PDF onto a mobile device or 2) print the PDF yourself.

  • Using a mobile device: I’ve found the PDF to be quite readable on my iPhone (in the Books app), although it helps that I’m near-sighted. It’s even easier to read on an iPad, other tablet, or laptop.
  • Printing the PDF: You can print the PDF yourself or at a local print shop. I highly recommend printing the 128 pages 2-sided to conserve paper and to reduce bulk and weight in the field. A comb or spiral binding, binder clip, or other binding will hold it together.

Your additions, corrections, comments, or questions are always welcome.

Cliffs above Bodie Creek at the state line

Copyright © Tim Messick 2024. All rights reserved.

Psathyrotes, Psathyrotopsis, and Trichoptilium (Oh, My!)

Recently I found myself looking closely at keys, descriptions, and photos of Psathyrotes annua (Annual turtleback) and P. ramosissima (“regular” Turtleback)—both rayless members of the Sunflower family, Asteraceae—to clear up some lingering uncertainty over how to distinguish the two. Then one taxon led to another and it became a larger project.

Psatyrotes annua is familiar to many who have wandered the perimeter of Mono Lake. It thrives in the sparsely vegetated sands and gravelly pumice of Mono’s prehistoric and post-1945 receding shorelines. It’s often misidentified as P. ramosissima, which so far, has probably not been found in the Mono Basin, but is common in the Mojave, Colorado, and western Sonoran Deserts. These two can be distinguished as follows:

  • Leaf blades generally clearly divided by deeply-set, anastomosing (cross-connecting) veins into irregular, ovate to polygonal areas (suggesting the segmented “scutes” of a turtle’s carapace); outer phyllaries ± wide, spatulate to obovate, widely spreading to reflexed; florets 21–26 (16–32) per head; pappus of 120–140 bristles in 3–4 series …… Psathyrotes ramosissima
  • Leaf blades generally weakly or not at all divided by veins into scute-like areas; outer phyllaries ± narrow, mostly lance-linear, ± erect to spreading; florets 13–16 (10–20) per head; pappus of 35–50 bristles in one series …… Psathyrotes annua
Psathyrotes ramosissima
Left: © Chloe & Trevor Van Loon/iNaturalist, Right: © Tom Chester/iNaturalist
Psathyrotes annua
Left: © Tim Messick/iNaturalist, Right: © Jim Morefield/iNaturalist

Exploring this small genus further, one finds there are currently three species in Psathyrotes, another close relative formerly in Psathyrotes now banished to Trichoptilium—a monotypic (one species) genus—and three more close relatives housed in Psathyrotopsis, a newer genus carved out of Psathyrotes.

Apparently no single key has been crafted that includes all 7 species of Psathyrotes, Trichoptilium, and Psathyrotopsis. The Jepson eFlora includes two Psathyrotes and the Trichoptilium. The Flora of North America (FNA) includes all three Psathyrotes, the Trichoptilium and one species of Psathyrotopsis, but not the other two, which appear to be narrow endemics in southern Coahuila, Mexico. Both Jepson and FNA make you work through ponderous keys for the entire sunflower family to distinguish these three genera. A paper on “Taxonomy of Psathyrotes” (Strother and Pilz 1975) includes a key to all of the taxa included in FNA, but not the third Psathyrotopsis, because it was described later (Turner 1993).

So, why not write a straightforward key that includes all seven taxa? Not so simple, it turns out, because existing keys don’t all use the same set of characters, and that newest species endemic to southern Coahuila (Psathyrotopsis hintoniorum) has never been included in a key with its relatives—and the paper describing it refers the reader to the formal description in Latin to figure out what distinguishes it from Psathyrotopsis purpusii. Thank goodness for Google Translate!

Below is an image of my key. (The very limited text formatting abilities in this WordPress blog don’t lend themselves to an indented key longer than 1 or 2 couplets.) The key is adapted from Baldwin 2012, Strother 2006a, Strother 2006b, Strother and Pilz 1975, and Turner 1973 (citations below). It has been modified from the originals by adding species, changing some terminology, rearranging some couplets, and adding some character states.

Here are photos of the rest of the species in Psathyrotes, Psathyrotopsis, and Trichoptilium (thank you to the iNaturalist contributors who allow their images to be used non-commercially!)

Psathyrotes pilifera Both: © slothiker/iNaturalist
Psathyrotyopsis hintoniorum
Left: © Arizona State University, Right: © New York Botanical Garden
(both are duplicate sheets of Guy Nesom 7648)
Psathyrotopsis purpusii Both: © José G. Flores Ventura/iNaturalist
Psathyrotopsis scaposa
Left: © Matt Reala/iNaturalist, Right: © Joey Santore/iNaturalist
Trichoptilium incisum
Left: © Jessica Irwin/iNaturalist, Right: © Irene/iNaturalist

And here are maps showing the known distribution of each species, produced (by me) using QGIS and Adobe Illustrator, based on occurrence data (the dots) from specimen locations served by the SEINet Portal Network and “Research Grade” observations from iNaturalist (note: some dots could be mismapped or misidentifications). Colored shading shows the approximate generalized distribution of each taxon.

References:
Baldwin, Bruce G. 2012. adapted from Strother (2006), Psathyrotes, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=571, accessed on November 28, 2022.
Strother, John L., 2006a. Psathyrotes in Flora of North America 21:416–418 (http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=126897), accessed on November 28, 2022
Strother, John L., 2006b. Psathyrotopsis in Flora of North America 21:364–365 (http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=126898), accessed on November 28, 2022.
Strother, John L and George Pilz. 1975. “TAXONOMY OF PSATHYROTES (COMPOSITAE: SENECIONEAE).” Madroño; a West American journal of botany 23, 24–40. https://www.biodiversitylibrary.org/partpdf/169365
Turner, B. L. 1993. “A new species of Psathyrotopsis (Asteraceae, Helenieae) from Coahuila, México.” Phytologia 75, 143–146. https://doi.org/10.5962/bhl.part.17305.


Copyright © Tim Messick 2024. All rights reserved.